Saturday, September 6, 2008
What are Corals?
Sorry for putting this post up so late. We should have placed this post on corals way earlier. So, just read on and you can find out more about those magnificent 'rainforest' of the sea.
Coral is a general term used to describe a group of cnidarians, which indicates the presence of skeletal material that is embedded in the living tissue or encloses the animal altogether. -National Oceanic and Atmospheric Administration, U.S. Dept. of Commerce. "Glossary of Coral Reef Terminology."
Corals themselves are tiny animals which belong to the group cnidaria (the "c" is silent). Other cnidarians include hydras, jellyfish, and sea anemones. Corals are sessile animals, meaning they are not mobile but stay fixed in one place.They feed by reaching out with tentacles to catch prey such as small fish and planktonic animals.
Corals are anthozoans, the largest class of organisms within the phylum Cnidaria. Comprising over 6,000 known species, anthozoans also include sea fans, sea pansies and anemones. Stony corals (scleractinians) make up the largest order of anthozoans, and are the group primarily responsible for laying the foundations of, and building up, reef structures. For the most part, scleractinians are colonial organisms composed of hundreds to hundreds of thousands of individuals, called polyps.
Corals live in colonies consisting of many individuals, each of which is called polyp. They secrete a hard calcium carbonate skeleton, which serves as a uniform base or substrate for the colony. The skeleton also provides protection, as the polyps can contract into the structure if predators approach. It is these hard skeletal structures that build up coral reefs over time. The calcium carbonate is secreted at the base of the polyps, so the living coral colony occurs at the surface of the skeletal structure, completely covering it. Calcium carbonate is continuously deposited by the living colony, adding to the size of the structure. Growth of these structures varies greatly, depending on the species of coral and environmental conditions-- ranging from 0.3 to 10 centimeters per year. Different species of coral build structures of various sizes and shapes ("brain corals," "fan corals," etc.), creating amazing diversity and complexity in the coral reef ecosystem. Various coral species tend to be segregated into characteristic zones on a reef, separated out by competition with other species and by environmental conditions.
Most corals are made up of hundreds of thousands individual polyps like this one. Many stony coral polyps range in size from one to three millimeters in diameter. Anatomically simple organisms, much of the polyp’s body is taken up by a stomach filled with digestive filaments. Open at only one end, the polyp takes in food and expels waste through its mouth. A ring of tentacles surrounding the mouth aids in capturing food, expelling waste and clearing away debris. Most food is captured with the help of special stinging cells called nematocysts which are inside the polyp' outer tissues, which is called the epidermis. Calcium carbonate is secreted by reef-building polyps and forms a protective cup called a calyx within which the polyps sits. The base of the calyx upon which the polyp sits is called the basal plate. The walls surrounding the calyx are called the theca. The coenosarc is a thin band of living tissue that connect individual polyps to one another and help make it a colonial organism.
As members of the phylum Cnidaria, corals have only a limited degree of organ development. Each polyp consists of three basic tissue layers: an outer epidermis, an inner layer of cells lining the gastrovascular cavity which acts as an internal space for digestion, and a layer called the mesoglea in between .
The diagram above shows the anatomy of a nematocyst cell and its “firing” sequence, from left to right. On the far left is a nematocyst inside its cellular capsule. The cell’s thread is coiled under pressure and wrapped around a stinging barb. When potential prey makes contact with the tentacles of a polyp, the nematocyst cell is stimulated. This causes a flap of tissue covering the nematocyst—the operculum—to fly open. The middle image shows the open operculum, the rapidly uncoiling thread and the emerging barb. On the far right is the fully extended cell. The barbs at the end of the nematocyst are designed to stick into the polyp’s victim and inject a poisonous liquid. When subdued, the polyp’s tentacles move the prey toward its mouth and the nematocysts recoil back into their capsules.
All coral polyps share two basic structural features with other members of their phylum. The first is a gastrovascular cavity that opens at only one end. At the opening to this cavity, commonly called the mouth, food is consumed and some waste products are expelled. A second feature all corals possess is a circle of tentacles, extensions of the body wall that surround the mouth. Tentacles help the coral to capture and ingest plankton for food, clear away debris from the mouth, and act as the animal’s primary means of defense.
While coral polyps have structurally simple body plans, they possess several distinctive cellular structures. One of these is called a cnidocyte—a type of cell unique to, and characteristic of, all cnidarians. Found throughout the tentacles and epidermis, cnidocytes contain organelles called cnidae, which include nematocysts, a type of stinging cell. Because nematocytes are capable of delivering powerful, often lethal toxins, they are essential to capturing prey, and facilitate coralline agonistic interactions.
Most corals, like other cnidarians, contain a symbiotic algae called zooxanthellae, within their gastrodermal cells. The coral provides the algae with a protected environment and the compounds necessary for photosynthesis. These include carbon dioxide, produced by coral respiration, and inorganic nutrients such as nitrates, and phosphates, which are metabolic waste products of the coral. In return, the algae produce oxygen and help the coral to remove wastes. Most importantly, they supply the coral with organic products of photosynthesis. These compounds, including glucose, glycerol, and amino acids, are utilized by the coral as building blocks in the manufacture of proteins, fats, and carbohydrates, as well as the synthesis of calcium carbonate (CaCO3). The mutual exchange of algal photosynthates and cnidarian metabolites is the key to the prodigious biological productivity and limestone-secreting capacity of reef building corals.
Zooxanthellae often are critical elements in the continuing health of reef-building corals. As much as 90% of the organic material they manufacture photosynthetically is transferred to the host coral tissue . If these algal cells are expelled by the polyps, which can occur if the colony undergoes prolonged physiological stress, the host may die shortly afterwards. The symbiotic zooxanthellae also confers its color to the polyp. If the zooxanthellae are expelled, the colony takes on a stark white appearance, which is commonly described as “coral bleaching”